Phylum: Charophyta   Family: Closteriacea

Closterium submoniliferum N.N. Woronichin 1924

First described by Woronichin (1924: 85) without illustration. Krieger (1935: 292, pl. 18: 10) made it a variety of Closterium moniliferum. Recently, Coesel, & Meesters (2023: 60) reinstated it as a full species. It differs from Cl. moniliferum in having smaller scattered pyenoids rather than large axile ones.
Cell dimensions: L: 350-425 µm; B: 58-79 µm; Ap: 18-28 µm; L/B: 5-6.
Infrequently found in circumneutral waters.

References: 
Coesel, P.F.M. & Meesters, K.J. (2023 second edition). Desmids of the Lowlands.
Krieger, W. (1935). Die Desmidiaceen Europas mit Berücksichtigung der aussereuropäischen Arten.
Van Westen, M. (2024). Sieralgen in Drenthe.
Woronichin, N.N. (1924). Algae nonnullae novae e Caucaso. VI.